IIS 7 Error - Collection is read-only

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http://serverfault.com – After copying an ASP.net MVC 3 application to another server I am getting the following error even though the IIS settings and web.config are identical to the previous site. I think it must be a permissions issue but the files look like they have the correct permissions, any ideas? Server Error in '/' Application. Collection is read-only. Description: An unhandled exception occurred during the execution of the current web request. Please review the stack trace for more information about the error and where it originated in the code. Exception Details: System.NotSupportedException: Coll (HowTos)