Hardware advice for video sharing website similar to Youtube but much smaller user base

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http://serverfault.com – Currently I have two servers which are named (ProcessServer1 and ProcessServer2) that will do nothing but convert videos that users upload, into flv or mp4 format. The conversion settings are 512k although I don't think this matters for what I'm asking since I believe it has to do with file sizes and input/output operations but we'll get to that in a minute. These two servers will then upload the videos to the video storage server (StorageServer1). StorageServer1's role is nothing but store the video files. MainServer1 is going to host the website (currently using a modified version of openso (HowTos)